Original reference as:
- Sato A & N. Takezaki, H. Tichy, F. Figueroa, W.E. Mayer & J. Klein. 2003. "Origin and speciation of haplochromine fishes in East African crater lakes investigated by the analysis of their mtDNA, Mhc genes, and SINEs". Molecular Biology and Evolution. vol. 20 (n. 9), pp. 1448–1462 (crc05436) (abstract)
Conservation: Haplochromis sp. 'lutotu' is not evaluated by the international union for the conservation of nature in the iucn red list of threatened species.